Revision of Sceliages Westwood, a millipede-eating genus of southern African dung beetles (Coleoptera : Scarabaeidae)

DOI
http://dx.doi.org/10.1071/IT01025
Publication Year
2002
Publication Site
Invertebrate Systematics
Journal Volume
16
Page Numbers
931–955
Family
Scarabaeidae
General topic
Taxonomy
Biodiversity/Biogeography
Specific topic
biotic interactions
Author

Forgie, Shaun A; Grebennikov, Vasily V; Scholtz, Clarke H.

Abstract Note

The genus <i>Sceliages</i> Westwood (Scarabaeinae : Scarabaeini) from southern Africa is revised. Seven species are recognised: <i>Sc. granulatus</i>, sp. nov. (Botswana, South Africa), <i>Sc. augias</i> Gillet (Angola, Democratic Republic of Congo, Zambia), <i>Sc. adamastor</i> (Le Peletier de Saint-Fargeau &#38; Serville) (South Africa), <i>Sc. brittoni</i> zur Strassen (South Africa), <i>Sc. difficilis</i> zur Strassen (South Africa, Zimbabwe), <i>Sc. gagates</i> Shipp (South Africa, Mo&#231;ambique) and <i>Sc. hippias</i> Westwood (South Africa). The new species is described and the others are redescribed. Neotypes are assigned to <i>Sc. adamastor</i> and <i>Sc. gagates</i>. A key to the species is provided and male genitalia and other diagnostic characters are illustrated. A phylogenetic analysis of the genus is presented. Distribution maps of all species are provided. Mature larvae of <i>Sc. hippias</i> are described, the first for the genus. They can be distinguished from other Scarabaeinae larvae by a markedly reduced torma on the epipharynx and complete absence of hypopharyngeal sclerites (oncyli). Millipede relocation and burial behaviour of the adults of <i>Sc. hippias</i> and <i>Sc. adamastor</i> are described. We also provide descriptions of the brood chamber and brood balls of <i>Sc. hippias</i>.