Finding a rare resource: Bornean Scarabaeoidea (Coleoptera) attracted by defensive secretions of Diplopoda

Scarab beetles of four species, Onthophagus penicillatus Harold, O. rudis Sharp, Phaeochridius derasus (Harold) and Phaeochroops gilleti (Benderitter) (Scarabaeoidea: Scarabaeidae and Hybosoridae) were attracted by quinonous defensive secretions of Harpagophoridae (Diplopoda) in Sabah, Borneo. Unlike the Afrotropics, where some specialists feed on fresh millipede carcasses, in Borneo only generalist necrophages use this resource. This may be caused by the low abundance of juliform Diplopoda in Borneo prohibiting specialization on this resource.