Adventive Onthophagus (Coleoptera: Scarabaeidae) in North America: Geographic ranges, diagnoses, and new distributional records

Five Old World species of the dung beetle genus Onthophagus (nuchicornis, depressus, bonasus, gazella, and taurus) are presently recorded in the United States, Canada, and Mexico. The current North American range of each, based on numerous literature records, specimens from private collections, from the insect collections of Cornell University and the Carnegie Museum of Natural History, and our own collecting, is mapped. A brief diagnosis, habitus photographs, and illustrations of other diagnostic features are provided to aid in accurate identification of these adventive species. We also report the presence of O. gazella in 2 new states in the U.S. (Kansas, Tennessee), and of O. taurus in 6 new states in the U.S. (Missouri, New York, Pennsylvania, Ohio, Tennessee, West Virginia).